Guangdong Provincial Key Laboratory for Plant Epigenetics, Longhua Bioindustry and Innovation Research Institute, College of Life Sciences and Oceanography, Shenzhen University, Guangdong Province, Shenzhen 518060, China. College of Physics and Optoelectronic Engineering, Shenzhen University, Guangdong Province, Shenzhen 518060, China. College of Horticulture Science and Engineering, Shandong Agricultural University, Tai'an, Shandong 271018, China. Department of Botany and Plant Sciences, Institute of Integrative Genome Biology, University of California, Riverside, California 92521, USA.